Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the division Selachimorpha[1] (or Selachii) and are the sister group to the Batoidea (rays and kin). Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Late Ordovician (458–444 million years ago).[2] The earliest confirmed modern sharks (selachimorphs) are known from the Early Jurassic around 200 million years ago, with the oldest known member being Agaleus, though records of true sharks may extend back as far as the Permian.[3] Sharks range in size from the small dwarf lanternshark (Etmopterus perryi), a deep sea species that is only 17 centimetres (6.7 in) in length, to the whale shark (Rhincodon typus), the largest fish in the world, which reaches approximately 12 metres (40 ft) in length.[4] They are found in all seas and are common to depths up to 2,000 metres (6,600 ft). They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river sharks, which can be found in both seawater and freshwater, and the Ganges shark, which lives only in freshwater.[5] Sharks have a covering of placoid scales (denticles) that protects the skin from damage and parasites in addition to improving their fluid dynamics. They have numerous sets of replaceable teeth.[6]
Several shark species are apex predators, which are organisms that are at the top of their food chain with select examples including the bull shark, tiger shark, great white shark, mako sharks, thresher sharks and hammerhead sharks. Some sharks are filter-feeding planktivores, such as the whale shark and basking shark, which are among the largest fish ever lived.
Sharks are caught by humans for shark meat or shark fins. Many shark populations are threatened by human activities. Since 1970, shark populations have been reduced by 71%, mostly from overfishing[7] and mutilating practice such as shark finning.[8][9] Until the 16th century,[10] sharks were known to mariners as "sea dogs".[11] This is still evidential in several species termed "dogfish", or the porbeagle.
The etymology of the word shark is uncertain. The most likely etymology states that the original sense of the word was that of "predator, one who preys on others" from the Dutch schurk, meaning 'villain, scoundrel' (cf. card shark, loan shark, etc.), which was later applied to the fish due to its predatory behaviour.[12]
A now disproven[original research?] theory is that it derives from the Yucatec Maya word xook (pronounced [ʃoːk]), meaning 'shark'.[13] Evidence for this etymology came from the Oxford English Dictionary, which notes that shark first came into use after Sir John Hawkins' sailors exhibited one in London in 1569 and posted "sharke" to refer to the large sharks of the Caribbean Sea. However, the Middle English Dictionary records an isolated occurrence of the word shark (referring to a sea fish) in a letter written by Thomas Beckington in 1442, which rules out a New World etymology.[14][original research?]